Prof Tamás Székely

Tamás Székely

Tamás Székely

Professor of Biodiversity
4 South 0.26

01225 383676 (tel)
01225 386779 (fax)


2007 – present Professor of Biodiversity, University of Bath
2000 – 2007     Lecturer, Senior Lecturer and Reader, University of Bath
1995 – 2000      Post-doctoral researcher, University of Bristol
1989 – 1995       Lecturer in Animal Behaviour, Kossuth University, Debrecen (Hungary)
1986 – 1989       Researcher, Hungarian Ornithological Society, Budapest (Hungary)
1983 – 1986       PhD in Animal Ecology, Kossuth University, Debrecen

2012 – 2013   Humboldt Research Award, University of Göttingen, Germany
2012 – 2013   Mercator-DFG Visiting Professor, University of Bielefeld, Germany
2011 – 2012    Erasmus Mundus Professor, University of Groningen, The Netherlands
2006 – 2007  Hrdy Visiting Fellowship, Harvard University, USA
2005 – 2006  Leverhulme Research Fellowship, University of Bath

I am an evolutionary biologist and my research focuses on sex roles, mating systems and parental care. I am particularly interested in field biology, phylogenetic comparative analyses and modelling.

My current research topics include:
• Behavioural ecology of mating systems, parental care and sexual selection
• Social behaviour especially birds and mammals
• Evolutionary ecology and phylogenetic comparisons and their applications
• Neuro-genomic bases of sex roles, mating and parenting
• Biodiversity conservation of wetland birds


Andras Liker, Gabriel Pena, Peter Long, Tamas Szekely, Richard James, Rene van Dijk (back row, from left to right), Araceli Tico, Freya Harrison, Fiona Burns, Ronald Jenner, Richard Young (middle), Clemens Kupper, Monif AlRashidi, Sama Zefania (front)

Post-doctoral researcher
Dr Kate Ashbrook – reintroduction of Great Bustard in the UK

PhD students
Cristina Carmona – Mating system, display behaviour and adult sex ratio in ploverS
Mohammed Al-Malki – Conservation of shorebirds in Saudi Arabia
Nina Ockendon – Social behaviour and genome evolution in birds
Alex Banks – Sexual conflict and sperm evolution in birds
Jorge Parra – Mating opportunities and breeding system of Malagasy plovers
Natalie dos Remedios – Phylogeography of plovers Charadrius spp.

Dr András Liker – Professor, University of Pannonia, Hungary
Dr Tatsuya Amano – Marie Curie Fellow, Cambridge University
Dr Sama Zefania – Scientific adviser, Malagasy National Parks, Madagascar
Dr Ákos Pogány – Post-doctoral researcher, Eötvös University, Hungary
Dr René van Dijk – Post-doctoral researcher, Sheffield University
Dr Peter Long – James Martin Fellow, Oxford University
Dr Clemens Küpper – Marie Curie Fellow, Sheffield University
Dr Mark O’Connell – Environmental consultant
Dr Freya Harrison – Research Fellow, Nottingham University
Dr Val Olson – Statistical consultant, London
Dr András Kosztolányi – Post-doctoral researcher, University of Debrecen
Dr Robert J Burnside – Post-doctoral research, University of East Anglia
Dr Araceli Argüelles-Tico – Post-doctoral researcher, Sheffield University
Dr Fiona Burns – Research scientist, Royal Society for Protection of Birds
Dr Monif Al-Rashidi – Dean of Scientific Research, Hail University, Saudi Arabia
Dr James St Clair – Post-doctoral researcher, St Andrews University
Dr Gabriel Pena – Conservation biologist, Mexico
Dr Martin Alejandro Serrano – Assistant Professor, University of Tlaxcala, Mexico
Dr István Szentirmai – Conservation biologist, Örség National Park, Hungary
Dr Fiona Sharpe – Environmental consultant
Dr Gavin Thomas – Lecturer, Sheffield University
Dr János Kis – Assistant Professor, Szent-István University, Budapest
Dr Zoltán Barta – Professor, University of Debrecen

I have ongoing collaborative projects with about one third of my former PhD students and postdocs. In addition, my recent/current collaborators include (in no particular order, see joint details in ResearchGate):

Prof Rob Freckleton, Sheffield University – phylogenetic comparative analyses
Prof Mike Bruford, Cardiff University – conservation genetics of Malagasy birds
Prof Terry Burke, Sheffield University – molecular ecology of shorebirds
Prof Innes Cuthill, Bristol University – parental care and sexual conflict in plovers
Prof John McNamara, Bristol University – game theoretic modelling
Prof Oliver Krüger, Bielefeld University – comparative analyses of brood parasitism
Dr Joe Hoffman, Biefeleld University – demographic analyses of Malagasy plovers
Prof Peter Kappeler, Göttingen University – sex ratio variation in mammals
Prof Jan Komdeur, University of Groningen – sexual conflict in penduline tits
Prof Franjo Weissing, University of Groningen – sex ratio evolution
Prof John Reynolds, Simon Fraser University – sexual conflict in shorebirds
Prof Bill Sutherland, Cambridge University – conservation biology of wetland birds
Prof Adám Miklósi, Eötvös University – social learning of sex roles
Prof Hans Hofmann, University of Texas – neuro-genomics of mating systems
Dr Lauren O’Connell, Harvard University – neuro-genomics of mating in plovers
Prof Larry Young, Emory University – sex roles in the bird brain
Dr Steve Dorus, Syracuse University – sperm competition in birds
Prof Robert Kelsh, University of Bath – evolution of melanin plumage in plovers
Dr Araxi Urrutia, University of Bath – social behaviour and comparative genomics
Dr Liu Yang, Sun-Yat Sen University – molecular ecology of plovers
Dr Pat Lee, Swansea University – phylogeography of plovers

These collaborative projects are funded by various national and international agencies including BBSRC, NERC, The Royal Society, The Leverhulme Trust, EU-Framework 6 & 7, LIFE+, German Science Foundation (DFG), Hungarian Research Fund (OTKA) and Mexican Scientific Fund (Conacyt).


Key Publications (see complete list of publications below)
1. Liker, A., R. P. Freckleton & T. Székely. 2013. The evolution of sex roles in birds is related to adult sex ratio. Nature Communications 4: 1587.
2. Moore, J. M, T. Székely, J. Büki, & T. J. DeVoogd. 2011. Motor pathway convergence predicts syllable repertoire size in oscine birds. Proceedings of National Academy of Sciences US 108: 16440-16445.
3. Székely, T., A. J. Moore & J. Komdeur (eds). 2010. Social behaviour: genes, ecology and evolution. Cambridge University Press, 1-562.
4. Fairbairn, D., W. Blanckenhorn & T. Székely (eds). 2007. Sex, size and gender roles. Evolutionary studies of sexual size dimorphism. Oxford University Press, 1-266.
5. Houston, A. I., T. Székely & J. M. McNamara. 2005. Conflict over parental care. Trends in Ecology and Evolution 20: 33-38.
6. Székely, T., R. P. Freckleton & J. D. Reynolds. 2004. Sexual selection explains Rensch’s rule of size dimorphism in shorebirds. Proceedings of National Academy of Sciences US 101: 12224 – 12227.
7. Blomqvist D., M. Andersson, C. Küpper, I. C. Cuthill, J. Kis, R. B. Lanctot, B. K. Sandercock, T. Székely, J. Wallander & B. Kempenaers. 2002. Genetic similarity between mates explains extra-pair parentage in three species of waders. Nature 419: 613-615.

I teach the following courses at Bath: Ecology and evolution (BB10012), Concepts in evolution (BB20040), Field course (BB20041), Data interpretation (BB20153), Behaviour and ecology (BB20173), Sexual conflict (BB30132/40193), Conservation biology (BB30156). In addition, I often contribute to postgraduate courses (PhD and Master courses) run by other universities, e.g. Groningen, Göttingen, Bielefeld and Debrecen.

Selected Recent Publications
1. Houston, A. I., T. Székely & J. M. McNamara. 2013. The parental investment models of Maynard Smith: a retrospective and prospective view. Animal Behaviour (invited review).
2. Moore, D., S. Wigby, S. English, S. Wong, T. Székely & F. Harrison. 2013. Selflessness is sexy: reported helping behaviour increases desirability of men and women as long-term sexual partners. BMC Evol Biol (in press).
3. Gamero, A., T. Székely & P. M. Kappeler. 2013. Delayed juvenile dispersal and monogamy, but no cooperative breeding in white-breasted mesites (Mesitornis variegata). Behavioral Ecology and Sociobiology (in press).
4. Székely, T., J. I. Hoffman & O. Krüger. 2013. Why do males care for their competitor’s offspring? Animal Behaviour Forum (in press).
5. Burns, F., N. McCulloch, N. dos Remedios, T. Székely, M. Bolton. 2013. Sex differences in behaviour but not mortality risk in a critically endangered shorebird. Ibis (in press).
6. Liker, A., R. P. Freckleton & T. Székely. 2013. The evolution of sex roles in birds is related to adult sex ratio. Nature Communications 4: 1587.
7. Williamson, L., M. Hudson, M. O’Connell, N. Davidson, R. Young, T. Amano & T. Székely. 2013. Areas of high diversity for the world’s inland-breeding waterbirds. Biodiversity and Conservation 22: 1501-1512.
8. Vincze, O., T. Székely, C. Küpper, M. AlRashidi, J. A. Amat, A. A. Ticó, D. Burgas, T. Burke, J. Cavitt, J. Figuerola, M. Shobrak, T. Montalvo & A. Kosztolányi. 2013. Local environment but not genetic differentiation influences biparental care in ten plover populations. PLoS ONE 8: e60998.
9. Moghadam, H. K., P. W. Harrison, G. Zachar, T. Székely & J. E. Mank. 2013. The plover neurotranscriptome assembly: transcriptomic analysis in an ecological model species without a reference genome. Molecular Ecology Resources (in press).
10. Ramm, S. A*, R. M. Jonker, K. Reinhold, T. Székely, F. Trillmich, T. Schmoll, H. Schielzeth, R. P. Freckleton. 2013. Comment on “Bateman in Nature: Predation on Offspring Reduces the Potential for Sexual Selection”. Science (in press) *All authors contributed equally to this work, and the author order was randomized.
11. Székely, T., V. Remeš, R. P Freckleton & A. Liker. 2013. Why care? Inferring the evolution of complex social behaviour. Journal of Evolutionary Biology 26: 1381-1391.
12. Burns, F., N. McCulloch, T. Székely & M. Bolton. 2013. The impact of introduced predators on an island endemic, the St. Helena Plover, Charadrius sanctaehelenae. Bird Conservation International (in press).
13. Zefania, S. & T. Székely. 2013. Madagascar plover Charadrius thoracicus. IN: R. Safford & F. Hawkins (eds). Birds of Madagascar. (in press)
14. Zefania, S. & T. Székely. 2013. White-fronted plover Charadrius marginatus. IN: R. Safford & F. Hawkins (eds). Birds of Madagascar. (in press)
15. Zefania, S. & T. Székely. 2013. Kittlitz’s plover Charadrius pecuarius. IN: R. Safford & F. Hawkins (eds). Birds of Madagascar. (in press)
16. Zefania, S. & T. Székely. 2013. Three-banded plover Charadrius tricollaris. IN: R. Safford & F. Hawkins (eds). Birds of Madagascar. (in press)

17. Garcia-Pena, G. E., D. Sol, A. N. Iwaniuk & T. Székely. 2012. Sexual selection on brain size in shorebirds (Charadriiformes). Journal of Evolutionary Biology 26: 878-888.
18. Burns, F., N. McCulloch, T. Székely & M. Bolton. 2013. More harm than good? Assessing the impact of nest exclosures for the endangered St. Helena Plover (Charadrius sanctaehelenae). Ibis (accepted).
19. Burnside, R.J., Zs. Végvári, R. James, S. Konyhás, G. Kovács & T. Székely. 2013. Human disturbance and conspecifics influence display site selection by Great Bustards Otis tarda. Bird Conservation International (accepted).
20. Küpper, C., S. V. Edwards, A. Kosztolányi, M. Alrashidi, T. Burke, P. Herrmann, A. Argüelles-Tico, J. A. Amat, M. Amezian, A. Rocha, H. Hötker, A. Ivanov, J. Chernicko & T. Székely. 2012. High gene flow on a continental scale in the polyandrous Kentish Plover Charadrius alexandrinus. Molecular Ecology 21, 5864–5879.
21. Székely, A. & T. Székely 2012. Sex ratio and the City. Dispatch. Current Biology 22: 684-685.
22. Pogány, Á. R. E. van Dijk, P. Horváth, T. Székely. 2012. Parental behavior and reproductive output in male-only and female-only cared clutches in the Eurasian penduline tit Remiz pendulinus. Auk 29: 773-781.
23. Pogány, Á., van Dijk, R.E., Menyhárt, O., Miklósi, Á., DeVoogd, T.J. T. Székely. 2012. Acoustic signaling in Eurasian Penduline Tits Remiz pendulinus: repertoire size signals male nest defence. Acta Zool Sci Hung (accepted).
24. Küpper, C., J. Augustin, S. Edwards, T. Székely, A. Kosztolányi, T. Burke, D. E. Janes. 2012. Triploid plover female provides support for a role of the W chromosome in avian sex determination. Biology Letters 8: 787-789.
25. Kosztolányi, A., C. Küpper, O. Chastel, C. Parenteauc, K. T. Yılmaz, Á. Miklósi, T. Székely, Á. Z. Lendvai. 2012. Prolactin stress response does not predict brood desertion in a polyandrous shorebird. Hormones and Behavior 61: 734-740.
26. van Dijk, R. E. J. Komdeur & T. Székely. 2012. Nest attendance does not predict offspring desertion by Eurasian penduline tit parents. Ethology (accepted)
27. Sutherland, W.J., J. A. Alves, T. Amano, C. H. Chang, N. C. Davidson, C. M. Finlayson, J. A. Gill, R. E. Gill, Jr., P. M. González, T. G. Gunnarsson, D. Kleijn, C. J. Spray, T. Székely, D. B.A. Thompson. 2012. An horizon scanning assessment of current and potential future threats facing migratory shorebirds. Ibis (in press).
28. AlRashidi, M., M. Shobrak & T. Székely. 2012. Integrating spatial data and nesting locations to predict the future impact of global warming on coastal habitats: A case study of shorebirds in Farasan Islands, Saudi Arabia. Saudi Journal of Biological Sciences (accepted).

29. Amano, T, T. Székely, K. Koyama, H. Amano & W. J. Sutherland. 2011. Addendum to ‘‘A framework for monitoring the status of populations: An example from wader populations in the East Asian-Australasian flyway’’. Biological Conservation, 143, 2238–2247. Biological Conservation 145: 278-295.
30. van Dijk, R. E., T. Székely, J. Komdeur, Á. Pogány, T. W. Fawcett & F. J. Weissing. 2012. Individual variation and the resolution of conflict over parental care in penduline tits. Proceedings of the Royal Society of London, B 279: 1927-1936.
31. Rheindt, F. E., T. Székely, S. V. Edwards, P. L. M. Lee, T. Burke, P. R. Kennerley, D. N. Bakewell, M. AlRashidi, A. Kosztolányi, M. A. Weston, W.-T. Liu, W.-P. Lei, Y. Shigeta, S. Javed, S. Zefania & C. Küpper. 2011. Conflict between genetic and phenotypic differentiation: the evolutionary history of a ‘lost and rediscovered’ shorebird. PLoS ONE 6, e26995.
32. Moore, J. M, T. Székely, J. Büki, & T. J. DeVoogd. 2011. Motor pathway convergence predicts syllable repertoire size in oscine birds. Proceedings of National Academy of Sciences, US 108: 16440-16445.
33. AlRashidi, M., A. Kosztolányi, M. Shobrak, C. Küpper & T. Székely. 2011. Parental cooperation in an extreme hot environment: natural behaviour and experimental evidence. Animal Behaviour 82: 235-243.
34. AlRashidi, M., A. Kosztolányi, M. Shobrak & T. Székely. 2011. Breeding ecology of the Kentish Plover Charadrius alexandrinus in Farasan Islands, Saudi Arabia (Aves: Charadriiformes). Zoology in the Middle East 53: 15-24.
35. Végvári, Zs, Z. Barta, P. Mustakallio & T. Székely. 2011. Consistent avoidance of human disturbance over large geographic distances by a migratory bird. Biological Letters 7: 814-817.
36. Bot, S., D. Brinkhuizen, Á. Pogány, T. Székely & R. van Dijk. 2011. Penduline tits in Eurasia: distribution, identification and systematic. Dutch Birding 33: 177-187.
37. Breitling, R., A Coleing, T. Peixoto, H. Nagle, G. Hancock, R. N Kelsh & T. Székely. 2011. An overview of spider fauna of Maio (Cape Verde Islands) with some additional records (Arachnida:Araneae). Zoologia Capoverdiana (in press).
38. Kosztolányi, A., Z. Barta, C. Küpper & T. Székely. 2011. Persistence of an extreme male-biased adult sex ratio in a natural population of polyandrous bird. Journal of Evolutionary Biology 24: 1842-1846.
39. Kelsh, R., T. Székely & S Stuart. 2011. Whys should biomedical scientists care about biodiversity? Current Biology 21: 210-211.
40. Burnside, R.J., I Carter, A. Dawes, D. Waters, L. Lock, P. Goriup & T. Székely 2011. The UK great bustard Otis tarda reintroduction trial. Oryx 46: 112-121.
41. AlRashidi, M., P. R. Long, M. O’Connell, M. Shobrak & T. Székely. 2011. Use of remote sensing to identify suitable breeding habitat for the Kentish plover and estimate population size along the western coast of Saudi Arabia. Wader Study Group Bulletin 53: 15-24.

42. Székely, T. & W. J. Sutherland. 2010. Hunting the cause of a population crash. Commentary. Nature 466: 448.
43. Remeš, V. & T. Székely. 2010. Domestic chickens defy Rensch’s rule: sexual size dimorphism in chicken breeds. Journal of Evolutionary Biology 23: 2754-2759.
44. Küpper, C., A. Kosztolányi, J. Augustin, D. A. Dawson, T. Burke & T. Székely. 2010. Heterozygosity-fitness correlations of conserved microsatellite markers in Kentish plovers Charadrius alexandrinus. Molecular Ecology 19: 5172–5185.
45. St Clair, J. J.H., G. Garcia-Pena, R. Woods, T. Székely. 2010. Presence of mammalian predators decreases tolerance to human disturbance in a breeding shorebird. Behavioral Ecology 21: 1285-1292.
46. St Clair, J. J. H, S. Poncet, D. K. Sheehan, T. Székely & G. M. Hilton. 2010. Responses of an island endemic invertebrate to rodent invasion and eradication. Animal Conservation 14: 66-73.
47. dos Remedios, N., P. L. M. Lee, T. Székely, D. A. Dawson & C. Küpper. 2010. Molecular sex-typing in shorebirds: a review of an essential method for research in evolution, ecology and conservation. International Wader Study Group Bulletin 117: 109-118 (invited review).
48. Amano, T, T. Székely, K. Koyama, H. Amano & W. J. Sutherland. 2010. A framework for monitoring the status of populations: an example from 3 wader populations in the East Asian-Australasian flyway. Biological Conservation 143: 2238-2247.
49. van Dijk R. E, Á. Pogány, J Komdeur, P Lloyd & T Székely. 2010. Sexual conflict predicts morphology and behavior in two species of penduline tits. BMC Evolutionary Biology 10: 107.
50. van Dijk, R. E., Mészáros, L., Velde, M., Székely, T., Pogány, Á., Szabad, J. & J. Komdeur 2010. Nest desertion is not predicted by cuckoldry in the Eurasian penduline tit. Behavioral Ecology & Sociobiology 64: 1425-1435.
51. Webb, T.J., V. A. Olson, T. Székely & R. P. Freckleton. 2010. Who cares? Quantifying the evolution of division of parental effort. Methods in Ecology & Evolution 1: 221-230.
52. Sol, D, N. Garcia, A Iwaniuk, K Davis, A Meade, A Boyle & T. Székely. 2010. Evolutionary divergence in brain size between migratory and resident birds. PLoS ONE 5: e9617.
53. St Clair, J. J. H., P. Herrmann, R. W. Woods & T. Székely. 2010. Female-biased incubation and strong diel sex-roles in the Two-banded Plover Charadrius falklandicus. Journal of Ornithology 151: 811-816.
54. Burns, F., T. Székely & M. Bolton. 2010. Leg flags versus colour rings: a comparison of marking methods using a small shorebird, the St Helena Wirebird. Wader Study Group Bulletin 117: 131-134.
55. van Dijk, R E., D. M. Brinkhuizen, T Székely & J. Komdeur. 2010. Parental care strategies in Eurasian penduline tit are not related to breeding densities and mating opportunities. Behaviour 147, Special Issue 1551-1565.
56. Moskát, Cs., M Bán, T Székely, J Komdeur, R W. G. Lucassen, L A van Boheemen & M E. Hauber. 2010. Discordancy or template-based recognition? Dissecting the cognitive basis of the rejection of foreign eggs in hosts of avian brood parasites. Journal of Experimental Biology 213: 1976-1983.
57. Brede, E G, P Long, S Zefania, M Rabenandrasana, T Székely & M Bruford. 2010. PCR primers for microsatellite loci in a Madagascan waterbird, the Sakalava rail (Amaurornis olivieri). Conservation Genetics Resources 2: 273-277.
58. AlRashidi, M, A Kosztolányi, C Küpper, I C Cuthill, S Javed & T Székely. 2010. The influence of a hot environment on parental cooperation of a ground-nesting shorebird, the Kentish plover Charadrius alexandrinus. Frontiers in Zoology 7:1.
59. St Clair, J. J. H., C Küpper, P Herrmann, R W. Woods & T Székely. 2010. Unusual incubation sex-roles in the rufous-chested dotterel Charadrius modestus. Ibis 152: 402-404.
60. Zefania, S., E. Razafimahatratra, P J Faria, M W Bruford, P R Long & T Székely. 2010. Morphometric and sexual size dimorphism in Malagasy plovers Charadrius spp. Ostrich 81: 173-178.
61. Székely, T., A. J. Moore & J. Komdeur (eds). 2010. Social behaviour: genes, ecology and evolution. Cambridge University Press, 1-562.
62. Székely, T., A. J. Moore & J. Komdeur. 2010. Introduction. IN Székely, T., A Moore & J Komdeur (eds). Social behaviour: genes, ecology and evolution. Cambridge University Press, 1-4.
63. Moore, A. J., Székely, T. & J. Komdeur. 2010. Prospects for research in social behaviour: systems biology meets behaviour. IN Székely, T., A. Moore & J. Komdeur (eds). Social behaviour: genes, ecology and evolution. Cambridge University Press, 538-550.
64. McGraw, L., T. Székely & L. J. Young. 2010. Pair bonds and parental behaviour. IN Székely, T., A. Moore & J. Komdeur (eds). Social behaviour: genes, ecology and evolution. Cambridge University Press, 271-301.

65. García-Peña, G. E., G. H. Thomas, J D. Reynolds & T. Székely. 2009. Breeding systems, climate and the evolution of migration in shorebirds. Behavioral Ecology 20: 1026-1033.
66. Harrison, F., Z. Barta, I C Cuthill & T. Székely. 2009. Conflict and cooperation between parents over care: a meta-analysis. Journal of Evolutionary Biology 22: 1800-1812.
67. Lislevand, T., J. Figuerola & T. Székely. 2009. Evolution of sexual size dimorphism in grouse and allies (Aves: Phasianidae) in relation to mating competition, fecundity demands and resource division. Journal of Evolutionary Biology 9: 1895-1905.
68. Küpper, C., J Augustin, A Kosztolányi, T Burke, J Figuerola and T. Székely. 2009. Kentish versus snowy plover: phenotypic and genetic analyses of Charadrius alexandrinus reveal divergence of Eurasian and American subspecies. Auk 126: 839-852.
69. Tjørve, K. M. C., G. E. García-Peña and T. Székely. 2009. Chick growth rates in Charadriides: comparative analyses of breeding climate, development mode and parental care. Journal of Avian Biology 40: 553-558.
70. Yu, M., Ming, M., Bao-wen, H., Brinkhuizen, D. & T. Székely. 2009. Nests and nest-site selection of white-crowned penduline tit Remiz coronatus in the Northern Xinjiang. Zoological Research 30: 565-570.
71. Olson, V. A., T. Webb, R. P. Freckleton & T. Székely. 2009. Are parental care trade-offs in shorebirds driven by parental investment or sexual selection? Journal of Evolutionary Biology 22: 672-682.
72. Kosztolányi A., I. C. Cuthill & T. Székely. 2009. Negotiation between parents over care: reversible compensation during incubation. Behavioral Ecology 20: 446-452.
73. Kosztolányi, A., S. Javed, C. Küpper, I. C. Cuthill, A. A. Shamsi & T. Székely. 2009. Breeding ecology of Kentish plover Charadrius alexandrinus in an extremely hot environment. Bird Study 56: 244-252.
74. Gergely Zs, L A. Mészáros, J Szabad & T Székely. 2009. Old nests are cues for suitable breeding sites in the Eurasian penduline tit, Remiz pendulinus. Journal of Avian Biology 40: 2-6.
75. Küpper, C., T Burke, T Székely & D A. Dawson. 2009. Enhanced cross-species utility of conserved microsatellite markers in shorebirds. BMC Genomics 9: 502.
76. Meininger P, T. Székely & D. Scott. 2009. Kentish plover Charadrius alexandrinus. IN: S. Delany, D. Scott, T. Dodman, D. Stroud (eds), An atlas of wader populations in Africa and Western Eurasia. Wetlands International, Wageningen, 229 – 235.

77. van Dijk, R & T Székely. 2008. Post-fertilization reproductive strategies. IN: Encyclopedia of Life Sciences, John Wiley & Sons, Ltd., Chichester [DOI: 10.1002/9780470015902.a0003665] (invited review).
78. van Dijk, R. E., Pilon, A. E., Szentirmai, I., Székely, T. & J. Komdeur. 2008 The influence of habitat structure on sexual conflict over care in penduline tits Remiz pendulinus. Ardea 96, 3-11
79. Pogány, Á., I. Szentirmai, J. Komdeur & T. Székely. 2008. Sexual conflict and consistency of offspring desertion in Eurasian penduline tit Remiz pendulinus. BMC Evolutionary Biology 8: 242.
80. Serrano-Meneses M. A., A Córdoba-Aguilar, M Azpilicueta-Amorín, E González-Soriano & T Székely. 2008. Sexual selection, sexual size dimorphism and Rensch’s rule in Odonata. Journal of Evolutionary Biology 21: 1259-1273.
81. Rabenandrasana, M. S. Zefania, P. Long, S T Seing, M C Virginie, M Randrianarisoa, R Safford & T Székely. 2009. Distribution, habitat and status of globally endangered endemic Sakalava rail of Madagascar. Bird Conservation International 19: 23-32.
82. van Dijk, R. E., J. Komdeur, M. van der Velde, I. Szentirmai, X. Yang, R. ffrench-Constant & T. Székely. 2008. Offspring sex ratio in the sequentially polygamous penduline tit Remiz pendulinus. Journal of Ornithology 149: 521-527.
83. Kingma, S., I. Szentirmai, T. Székely, V. Bokony, M. Bleeker, A. Liker, J. Komdeur. 2008. Sexual selection and the function of a melanin-based plumage ornament in promiscuous penduline tits Remiz pendulinus. Behavioral Ecology and Sociobiology 62: 1277-1288.
84. Long, P. R., S. Zefania, R. H. ffrench-Constant & T Székely. 2008. Estimating population size of an endangered shorebird, the Madagascar plover Charadrius thoracicus, using a habitat suitability model. Animal Conservation 11: 118-127.
85. Zefania, S., R. ffrench-Constant, P. R. Long & T. Székely. 2008. Breeding distribution and ecology of the endangered Madagascar plover Charadrius thoracicus. Ostrich 79: 43-51.
86. Olson, V.A., Liker A., Freckleton R. P. & T. Székely. 2008. Parental conflict in birds: comparative analyses of offspring development, ecology and mating opportunities. Proc Roy Soc. London B 275: 301-307.
87. Serrano-Meneses, M. A., A. Córdoba-Aguilar & T. Székely. 2008. Sexual size dimorphism: patterns and processes. IN: Córdoba-Aguilar, A. (ed.) Dragonflies and damselflies: model organisms for ecological and evolutionary research. Oxford University Press, Oxford, 231-248.
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